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the relationship between the видовыми characteristics and properties of the environment typically непрямо by двухшагового analysis. first, the abundance of species is associated with environmental conditions, and the reaction of the variability properties of the environment compares then with biological or physiological characteristics of the species (Thuiller et al., 2004; Santoul et al., 2005; Brind "amour et al., 2009). analysis of RQL allows correlating the ecological characteristics of species to environmental conditions (Doledec et al. 1996). this analysis explores the joint structure between the three tables of data: r - table (which contains the variables of the environment), the q table (which contains the species characteristics) and the table (all types) (Doledec et al., 1996; Dray et al., 2002). the table serves as a connection between tables r and q and measures the intensity of the connection between them. before the actual analysis, there are three separate analysis. correspondence analysis applied to the matrix, which are the optimal корреляционную structure between sites, and the balance of species. ordination of tables r and q is performed through the analysis of the main component. thus, the RQL performs analysis of the коинерции cross matrices r, q and l analysis. this maximizes the ковариацию between scale of sites, taking into account the characteristics of the environment, expressed as the r, and the balance of species according to their environmental properties expressed by the matrix q (minden et al., 2012). the result could be obtained by the combination of the best coordination sites on their characteristics of the environment, coordination of their properties and, at the same time, ordination of species and sites (Thuiller et al., 2006). RQL analysis consists of three separate ординационных decision with the maximization of ковариации between feature types and properties of the environment through the analysis of коинерции (bernhardt - Romermann et al., 2008). further, the hierarchical clustering analysis of the weights of the two axes RQL varda method provides functional group (minden et al., 2012). the optimal number of groups can be obtained by the criterion of калинского (Calinski, Harabasz, 1974). clusters show the distribution of species in the space of particular species, eco - logical space (minden et al., 2012).
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